I'm late to this (apologies), but just a couple of thoughts.
First thing that I haven't seen mentioned (I may have missed it though), have you checked the HT fuse? (If it is blown, replacing won't solve the issue that caused it to blow). We know the main fuse is ok because the filaments light up.
Question, do "all" filaments light up? There are two filaments in each pre-amp tube.
You saw light flashes coinciding with audible popping sound through speakers. That you saw the flashing would (most likely) seem to indicate that the light source was external to the amp's chassis. Possibly a tube arcing internally. Tubes, both pre-amp & power tubes can arc (in different ways) & still "look" ok. It should result in a blown HT, or main fuse though. This would mean "no sound" at all.
Check the HT fuse & get back to us, we'll go from there. Cheers