Why my 1959 Fender Concert amp is running 507 VDC on the plates should be 456 VDC
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The bias is too cold.
When the bias is adjusted to around 70% the plate voltage will drop down.
Another couple things is that the screen resistors can be bad, or the speaker impedance will be incorrect.
Those 2 things will also affect the power supply voltages.
Without the proper load on the power supply, the voltage will climb up. That's ohm's law.
Cold bias means less load on the power supply, so higher voltage would be expected.
True, 110 volt or 115 volt primary running on 120 volts can affect the plate volts by a few percent.
But that alone does not explain the 50 volt discrepancy. Try the bias adjust first...
If you do not have a bias meter or a scope you should get something.
"Dramatic change in tone..."
Um....not really.
But we should keep in mind that 500+ volts DC is a bit high.